"It was actually a very sad sight to see all those dead elephants, especially one of the dead females who had a very young calf of about three months old. The calf was trying to wake the dead mother up," he said.

The WWF estimates that there are fewer than 1,500 Borneo pygmy elephants in the wild, most of them in Sabah state.

They are the smallest elephant subspecies, with babyish faces, long tails and straight tusks. They are threatened by logging, hunting and increasing contact with humans.

Mr Masidi said the death of "these majestic and severely endangered Bornean elephants is a great loss to the state".

"If indeed these poor elephants were maliciously poisoned, I would personally make sure that the culprits would be brought to justice and pay for their crime," he said.

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